Related Clues:

• No bones back there - and no conifers, either, in the end
• No backbone and no conifers at last
• Make no bones about it: there are no conifers like this below the south
• No trees like this at last for the invertebrates
• Is the South so weak as to be devoid of some conifers?
• 'The South has no backbone and no conifers, either (9)'
• Vacillating invertebrate
• Lacking (moral) courage
• Completely lacking courage
• Not like Mrs Tiggy-Winkle to be so lily-livered!
• Weak; invertebrate
• Weakened by lack of a column?
• Old Bob sleeps in? Terribly weak!
• Invertebrate
• Yellow roses are not